Koba

Koba is a secondary character in Rise of the Planet of the Apes and the main antagonist of Dawn of the Planet of the Apes. He is a bonobo that was used as a test subject in Gen-Sys Laboratories, causing him to develop a hatred against humans. He later joined Caesar's ape colony and became his adviser and second-in-command.

Background
In his younger years, Koba lived with his mother in an ape facility, and was taught American Sign Language. After Koba's mother was beaten to death by their caregiver, Roger, Koba was sold to a TV studio run by a man named Tommy. There, Koba became aligned with another ape, Milo, and the two were taught to perform tricks for the show and were electrocuted if they failed. After low ratings resulted in the show being cancelled, a drunken Tommy took his anger out on Koba and Milo, beating them and blinding Koba in one eye with a lit cigarette. Tommy later committed suicide and Koba was captured by animal control officers and separated from Milo, being moved to several laboratories, during which Koba developed self-harming tendencies. At one point, Koba formed a bond with Gen-Sys neuroscientist Amol, who used Koba for several experiments. However, Amol's superior, Steven Jacobs fired him and confronted Koba, deriding him as an animal. Koba, finally reaching his breaking point, developed a hatred against humanity, focusing on Jacobs specifically as the source of his pain.

Rise of the Planet of the Apes
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Dawn of the Planet of the Apes
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Trivia

 * Koba is the first bonobo to appear in the Planet of the Apes series. Ironically, bonobos in real-life are generally friendly and gentle, whereas Koba is the most vicious ape in the Ape Village.
 * Koba is named after Joseph Stalin, who used Koba as an alias before the Russian Revolution.
 * At 26, Koba is the oldest known ape in the series.